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\textbf{Gustaf Granath}$^\star$, \footnote{\textit{Gustaf [email protected]}} {Department of Ecology, Swedish University of Agricultural Sciences Box 7044, SE-750 07 Uppsala, Sweden.}'\\  \textbf{Christopher D. Evans}, {Centre for Ecology and Hydrology, Bangor, UK.}\\  \textbf{Anna Landahl}, {Department of Aquatic Sciences and Assessment, Swedish University of Agricultural Sciences Box 7050, SE-750 07 Uppsala, Sweden.} \\  \textbf{Jens Fönster}, {Department of Aquatic Sciences and Assessment, Swedish University of Agricultural Sciences Box 7050, SE-750 07 Uppsala, Sweden.}\\  \textbf{Stephan Kölhler}, {Department of Aquatic Sciences and Assessment, Swedish University of Agricultural Sciences Box 7050, SE-750 07 Uppsala, Sweden.} \\