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Gustaf ^{1,2*}Gustaf  Granath Anna ^{2}Anna  Landahl Jens ^{2}Jens  Fönster Stephan ^{2}Stephan  Kölhler ^{1}Department of Ecology, Swedish University of Agricultural Sciences Box 7044, SE-750 07 Uppsala, Sweden.  ^{2}Department of Aquatic Sciences and Assessment, Swedish University of Agricultural Sciences Box 7050, SE-750 07 Uppsala, Sweden.  *Corresponding author: [email protected]