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Gustaf Granath\textsuperscript{1,2*}, Anna Landahl\textsuperscript{2}, Jens Fönster\textsuperscript{2}, Stephan Kölhler\textsuperscript{2}  \begin{enumerate}  \item Department of Ecology, Swedish University of Agricultural Sciences Box 7044, SE-750 07 Uppsala, Sweden.  \item Department of Aquatic Sciences and Assessment, Swedish University of Agricultural Sciences Box 7050, SE-750 07 Uppsala, Sweden.  \end{enumerate}