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Although this model neatly explains the increased silicate weathering during the rise of ectomycorrhizal fungi over the past 120 million years, we have major issues with the reality of the assumptions made.  The main part of protons exuded by plant roots and mycorrhizal fungi is in exchange for nutrients (cations and NH4+).   Most of these nutrient come from organic matter breakdown.  Ergo, Then  most proton are exuded where these nutrients are taken up. That is mostly at CEC sites (clays and organic matter) or close to the degrading organic matter itself, and not specifically close to nutrient-bearing minerals!  Fungal length measurements on individual grains from the top soil in a boreal pine forest does show a preferential colonization of feldspars over quartz, but as quartz was the dominant mineral, most fungal length was still found on the quartz grains.  Also, mineral mesh bag incubation studies do not show a clear effect of incubated mineral type on fungal colonization rates\cite{Rosenstock_2016}. These observations undermine the validity of the assumption that all ectomycorrhizal exchange processes take place only at nutrient-bearing minerals.