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\textbf{Pumping at variable food levels.} All panels correspond to recordings from wild-type animals. animals at $4x$ magnification.  In panels A-D each animal was assayed for $30$ min in the presence of a constant concentration of food (\textit{E. coli} OP50). In panels E-I each animal was assayed for $60$ min in and bacterial food concentration oscillated between high ($OD_{600} = 4.0$) and low ($OD_{600} = 0.0$) values with a period of $300$ sec. (A) Distributions of wild-type instantaneous pumping rates at a food concentration of $OD_{600} = 4.0$ were affected by the flow rate in the device. Flow rates of $5$ $\mu l / min$ and $200$ $\mu l / min$ are plotted in red and blue, respectively. $N_{5 \  \mu l / min} = ???$, $N_{200 \  \mu l / min} = ???$ animals (B) The distribution of instantaneous pumping rates in the absence of bacterial food ($OD_{600} = 0.0$).Data shown in panels A-B was acquired for $30$ min per animal at $4x$ magnification.  (C-D) Mean pumping rates and duty ratios for the data presented in panels A-B. The duty ratio was defined as the fraction of time occupied by continuous pumping, i.e., when consecutive pumps are up to 250 were $\leq 250$  ms apart. (E) Representative pumping dynamics in the presence of oscillating food availability. The concentration of food flowing through the device was oscillated between $OD_{600} = 4.0$ and $0$ with a period of 360 sec at a flow rate of 200 $\mu l / min$. (F) The mean pumping rates and food levels averaged over the corresponding nine stimuli shown for each of the individual animals in panel E. (G) Mean food levels averaged over stimuli and animals. Shaded areas in panels F-G denote $\pm$ s.e.m. (H) Response-triggered average to the periodic stimuli. For each incidence of the pumping rate exceeding $2$ Hz, the preceding $180$ sec of data were averaged. Grey lines depict data from individual animals and the blue line depicts the population mean. (I) Fraction of responses per animal. A response to a stimulus was defined as an increase in pumping rate during high food levels that was at least $2$ s.e.m higher than the mean rate at the preceding $OD_{600} = 0.0$ period. \label{fig:results}