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\textbf{Pumping at variable food levels.} All panels correspond to recordings from wild-type animals at $4x$ magnification. In panels A-D each animal was assayed for $30$ min in the presence of a constant concentration of food (\textit{E. coli} OP50). In panels E-I each animal was assayed for $60$ min. Bacterial food flowed at a rate of $200 \ \mu l / min$ and oscillated between high ($OD_{600} = 4.0$) and low ($OD_{600} = 0.0$) levels with a period of $300$ $360$  sec. (A) Distributions of wild-type instantaneous pumping rates at a food concentration of $OD_{600} = 4.0$ were affected by the flow rate in the device. Flow rates of $5$ $\mu l / min$ and $200$ $\mu l / min$ are plotted in red and blue, respectively. $N_{5 \ \mu l / min} = ???$, $N_{200 \ \mu l / min} = ???$ animals (B) The distribution of instantaneous pumping rates in the absence of bacterial food ($OD_{600} = 0.0$). $N = ???$ (C-D) Mean pumping rates and duty ratios for the data presented in panels A-B. The duty ratio was defined as the fraction of time occupied by continuous pumping, i.e., when consecutive pumps were $\leq 250$ ms apart. $N_{OD=0} = ???$, $N_{OD=2} = ???$, $N_{OD=4, \ 5 \mu l / min} = ???$, $N_{OD=4, \ 200 \mu l / min} = ???$. (E) Left: representative examples of pumping dynamics of individual animals in the presence of oscillating food availability. Right: the mean pumping rates and food levels per animal, i.e., data were averaged over the corresponding nine stimuli for each of the animals on the left. (F-G) Mean pumping rates and food levels averaged over stimuli and all animals. Shaded areas denote $\pm$ s.e.m. (H) Response-triggered average to the periodic stimuli. For each incidence of the pumping rate exceeding $2$ Hz, the preceding $180$ sec of data were averaged. Grey lines depict data from individual animals and the blue line depicts the population mean. (I) Fraction of responses per animal. A response to a stimulus was defined as an increase in pumping rate during high food levels that was at least $2$ s.e.m higher than the mean rate at the preceding $OD_{600} = 0.0$ period. In panels F-I $N = 9???$ animals. \label{fig:results}