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the chemical composition of freshly degrading plant biomass. The same substrate mixture was added to microcosms in each series, however, for each series except the control, one  substrate was substituted for its $^{13}$C counterpart; $^{13}$C-cellulose in one series, $^{13}$C-xylose in another, and no $^{13}$C-labeled substrates in the control. Microcosm amendments are shorthand identified in the following figures by the following code: "13CXPS" refers to the amendment with $^{13}$C-xylose (that is $^{13}$\textbf{C} \textbf{X}ylose  \textbf{P}lant \textbf{S}imulant), "13CCPS" refers to the $^{13}$C-cellulose amendment and "12CCPS" refers to the amendment that only contained $^{12}$C substrates (i.e. control). Xylose or cellulose were chosen to carry the isotopic label to contrast C assimilation for labile, soluble C (xylose) versus insoluble, polymeric C (cellulose). 5.3 mg of C substrate mixture per gram soil was added to each microcosm representing 18\% of the total soil C. The mixture included  0.42 mg xylose-C and 0.88 mg cellulose-C g soil$^{-1}$. Microcosms were harvested at days 1, 3, 7, 14 and 30 during a 30 day incubation. $^{13}$C-xylose assimilation peaked immediately and tapered over the 30 day incubation whereas $^{13}$C-cellulose assimilation peakedat  two weeks of (Figure~\ref{fig:ord}). after amendment additions(Figure~\ref{fig:ord}).  % Fakesubsubsection:We sequenced SSU rRNA gene amplicons from a  We sequenced SSU rRNA gene amplicons from a total of 277 CsCl gradient