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length can demonstrate patterns in substrate use specificity, and 3) $^{13}$C  incorporation can be established at high resolution taxonomic groups such  as 97\% sequence identity OTUs.  Microorganisms sequester atmospheric carbon and respire soil organic matter  (SOM) influencing climate change on a global scale but which microbial lineages  transform different soil C components are not established. Molecular tools will  unravel the soil microbial food web and reveal how specific microbial lineages  impact soil C flux. We present a cultivation independent, molecular, DNA-SIP  method to chart C use into microbial lineages. Our results show physiologically  undefined cosmopolitan microbial lineages decompose cellulose. We also show  phylogenetic groups rise and fall and are supplanted by others in activity over  7 days in response to labile C addition.   % Fakesubsubsection:HR-SIP allows us to assess C substrate specificity  HR-SIP allows us to assess C substrate specificity and temporal dynamics of  $^{13}$C-incorporation. C substrate specificity is assessed by measuring the BD 
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Soluble C more robust to temperature changes because or redundancy? But we  found similar numbers of xylose and cellulose degrader  Microorganisms sequester atmospheric carbon and respire soil organic matter  (SOM) influencing climate change on a global scale but which microbial lineages  transform different soil C components are not established. Molecular tools will  unravel the soil microbial food web and reveal how specific microbial lineages  impact soil C flux. We present a cultivation independent, molecular, DNA-SIP  method to chart C use into microbial lineages. Our results show physiologically  undefined cosmopolitan microbial lineages decompose cellulose. We also show  phylogenetic groups rise and fall and are supplanted by others in activity over  7 days in response to labile C addition.  \subsection{Ecological strategies of xylose and cellulose degrading OTUs in  soil}