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in soils despite the significant contribution of microbial activity to global  soil C flux. \citep{Allison2010,Six2006,Treseder2011}. It remains difficult to  predict the response of soil C to changes in climate  \citep{Nett_2001,Davidson2006a,Todd-Brown2013} \citep{Neff_2001,Davidson2006a,Todd-Brown2013}  but predictions of climate change feedbacks on soil C flux improve when biogeochemical models explicitly  represent microbial physiology \citep{Wieder2013}. However, we still know  little of the ecophysiology of soil microorganisms. Efforts that describe soil