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approach that expands upon previous nucleic acid SIP methods in three dimensions:  1) temporally, we sample isotopically labeled substrate amended microcosms at multiple   time points; 2) spacially, we assay more fractions along the CsCl gradients; and 3),   bioinformatically, we interrogate taxa at the level or of  OTU for isotope incorporation employing cutting edge statistics for assessing differential abundance in microbiome  datasets.