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\subsection{Cellulose degraders exhibit higher substrate specificty than xylose  utilizers} Cellulose responders exhibited a greater shift in BD than xylose  responders in response to isotope incorporation (Figure~\ref{fig:shift},  p-value 1.86e$^{-06}$). $^{13}$C-cellulose responders shifted on average 0.0163  g/mL (sd 0.0094) whereas xylose responders shifted on average 0.0097 (sd  0.0094). For reference, 100\% $^{13}$C DNA shifts X.XX g/mL relative to the BD  of its $^{12}$C counterpart. DNA BD increases as its ratio of $^{13}$C to  $^{12}$C increases. An organism that only assimilates C into DNA from a  $^{13}$C isotopically labeled source, will have a greater $^{13}$C:$^{12}$C  ratio in its DNA than an organism utilizing a mixture of isotopically labeled  and unlabeled C sources. Upon labeling, DNA from an organism that incorporates  exclusively $^{13}$C will increase in buoyant density more than DNA from an  organism that does not exclusively utilize isotopically labeled C. Therefore  the magnitude DNA buoyant density shifts indicate substrate specificity given  our experimental design as only one substrate was labeled in each amendment. We  measured density shift as the change in an OTU's density profile center of mass  between corresponding contol and labeled gradients. Density shifts, however,  should not be evaluated on an individual OTU basis as a small number of density  shifts are observed for each OTU and the variance of the density shift metric  at the level of individual OTUs is unknown. It is therefore more informative to  compare density shifts among substrate responder groups. Further, density  shifts are based on relative abundance profiles and would be theoretically  muted in comparison to density shifts based on absolute abundance profiles and  should be interpreted with this transformation in mind. It should also be noted  that there was overlap in observed density shifts between $^{13}$C-cellulose  and $^{13}$C-xylose responder groups suggesting that although cellulose  degraders are generally more substrate specific than xylose utilizers, some  cellulose degraders show less substrate specificity for cellulose than some  xylose utilizers for xylose (Figure~\ref{fig:shift}), and, each responder group  exhibits a range of substrate specificites (Figure~\ref{fig:shift}).  diff --git a/layout.md b/layout.md  index 76d9d2f..9e09d80 100644  --- a/layout.md  +++ b/layout.md 
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Significance.tex  Introduction.tex  Results.tex  BD.tex  rrn.tex  Discussion.tex  ecological_strategies.tex