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Our study is consistent with carbon degradative succession that has previously been demonstrated \cite{Bastian_2009} (more refs). We demonstrate a rapid decrease in the labile carbon, xylose, confirmed by its \textsuperscript{13}C label incorporation into the microbial community DNA during the first 7 days of the experiment, after which, the label is not detectable in the DNA. Subsequently our data demonstrates a slow degradation of the more recalcitrant, polymeric carbon demonstrated by \textsuperscript{13}C-cellulose label incorporation into the microbial community DNA at 14 and 30 days. We did not observe the \textsuperscript{13}C-cellulose signal leave the DNA within the time limits, 30 days, of our experiment. This degradative succession is also confirmed by isotopic analysis of the soil from the microcosms (Table S1). NMDS demonstrates microbial succession and based on xylose and cellulose treatments separating away from the control, but in opposing directions indicating different microbial community members are responsible for degradation of the two C substrates.   We did not observe consistent C utilization at the phylum level although both xylose and cellulose utilization were observed across 7 phyla each revealing a high diversity of bacteria able to utilize these substrates. The high taxonomic diversity may enable substrate metabolism under a broad range of environmental conditions \cite{Goldfarb_2011}. Other studies of microbial communities have observed a positive correlation with taxonomic or phylogenetic diversity and functional diversity \cite{Fierer_2012,Fierer_2013,Philippot_2010,Tringe_2005,Gilbert_2010,Bryant_2012}. A study on pre-agricultural prairie soils observed a decrease in functional capabilities of soil microbial communities with decreasing taxonomic diversity suggesting they do not exhibit a high degree of functional redundancy \cite{Fierer_2013}. Our Specific functional attributes are shared among diverse, yet distinct, taxa while closely related taxa may have very different physiologies and environmental tolerances (14). "\cite{Fierer_2012}Our  findings, along with similar studies, add to the growing collection of data suggesting that community membership matters. is importance to biogeochemical processes.  This calls for a need to examine substrate utilization by discrete microbial taxa within whole community studies, versus culture isolation, to better understand how specific community members function within the whole. distinct taxa can share specific functional attributes and closely related taxa may have very different physiologies and environmental tolerances (14). "\cite{Fierer_2012} SIP-NGS