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non-DNA-sequencing microbial community prifiling methods such as DGGE CITE and  tRFLP CITE, and currently to large experiments with multiple isotopically   labelled substrates coupled to high-throughput amplicon and   shotgun DNA sequencing \citep{Verastegui_2014}. We present a high-resoltion high-resolution  nucleic acid SIP (HR-SIP)   approach that expands upon previous nucleic acid SIP methods in three dimensions:  1) temporally, we sample isotopically labeled substrate amended microcosms at multiple