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HR-SIP allows us to assess C substrate specificity and temporal dynamics of  $^{13}$C-incorporation. C substrate specificity is assessed by measuring the BD  shift of OTU DNA upon $^{13}$C incorporation. OTUs that incorporate more  $^{13}$C per unit DNA have greater specify specificity  for the labeled substrate than OTUs that incorporate less $^{13}$C per unit DNA. $^{13}$C-cellulose incorporating  OTUs as a group displayed greater substrate specificity than $^{13}$C-xylose  incorporating OTUs. This suggests that polymeric C-degraders tend to be