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  **dicuss besemer/battin study**    From Besemer et al. 2012 "Besemer et al. (2007) compared community succession in  stream biofilms and found consistent differences between the biofilm and stream water communities, which indicate the existence of a specific biofilm community. On the basis of these previous findings, we hypothesize that the assembly of a local biofilm community is not a mere reflection of the source community suspended in the overlying stream water.  We are aware that niche diversification could, nevertheless, support a high diversity in biofilms (Jackson et al., 2001; Besemer et al., 2007).  Besemer et al. 2012 integrated plankton samples overtime before sequencing  reconcile with Battin’s work   
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In summary this study shows mechanistic links between large scale community level dynamics the underlying populations level that drive them. Ulimately large scale changes in ecosystem processes are driven by composite effects of microbial communities actiing as a synthesis of physiological events embedded in a complex biotic and abiotic matrix.   This resulst is consistent with carbon subsidies affecting the heterotrophic but not the autotrophic community and the similar lack of a shift in algal abundance within the biofilm of the autotroph community.