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Biofilm libraries were more even in shape than planktonic libraries (Figure 9) and OTUs enriched in biofilm libraries were enriched less dramatically than planktonic enriched OTUs. It's possible the planktonic environment is more uniform with respect to niche space and therefore produces more skewed rank abundance distributions. Similarly to the richness results, we found the shape contrasts of rank abundance distributions between biofilm and planktonic libraries in our data to be in contrast to that reported by \citet{22237539} although this may be an artifact of each study's different experimental design (see above).   \subsection{Enriched Taxa in High Carbon (C:P = 500) Treatment}  OTUs enriched in the high carbon amended mesocosm (C:P = 500) included classic copiotroph genera such as  In summary this study shows mechanistic links between large scale community level dynamics the underlying populations level that drive them. Ulimately large scale changes in ecosystem processes are driven by composite effects of microbial communities actiing as a synthesis of physiological events embedded in a complex biotic and abiotic matrix.