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\section{Acknowledgements}  This research was conducted as part of the 2010 Marine Biological Laboratory’s Microbial Diversity Course. Funding was supplied by the MBL, the Bernard Davis Endowed Scholarship Fund, and the Selman A. Waksman Endowed Scholarship. EKH would like to thank Dan Buckley and Steve Zinder for organizing the course, Marshall Otter, Mathew Erickson and Erickson,  Hugh Ducklow and Jay T. Lennon for analytical support, support and the  Austrian FWF MICDIF award to Tom Battin for salary support.