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explain the merging of the planktonic and biofilm bacterial membership in the  highest C treatment.   %\subsection{Lifestyle \subsection{Lifestyle  (biofilm or planktonic) Enriched OTUs} %  %There There  are only a few studies that attempt to compare biofilm community %composition composition  and the overlying planktonic community \citep{Besemer_2007, %22237539, 22237539,  Jackson_2001, Lyautey_2005}. Those studies illustrate community %composition composition  among the two habitats are unique with very few taxa found in both. %This This  is consistent with our findings in this experimental system with a natural %marine marine  planktonic source community. In addition, our study also evaluated %algal algal  community composition which showed a similar result suggesting that both %the the  algal and bacterial biofilm communities form from phylogenetically unique %organisms organisms  that exist in low abundance in surrounding habitat (i.e. the %plankton) plankton)  but are readily enriched in the biofilm lifestyle. Most of the %biofilm biofilm  enriched algal OTUs were \textit{Bacillariophyta} although there were %also also  many \textit{Bacillariophya} OTUs enriched in the planktonic libraries. We %also also  found \textit{Cryptophyta} and \textit{Viridiplantae} were more uniformly %enriched enriched  in the planktonic algal libraries. It appears that these broad %taxonomic taxonomic  groups were selected against in biofilms under our experimental %conditions. conditions.  Bacterial OTUs enriched in planktonic samples displayed more %dramatic dramatic  differential abundance patterns than bacterial OTUs enriched in %biofilm biofilm  samples, but, biofilm enriched bacterial OTUs were spread across a %greater greater  phylogenetic breadth (Figure~\ref{fig:l2fc}). This is also consistent %with with  the idea of greater niche diversity in the biofilm environment as opposed %to to  the plankton. Greater niche diversity should select for a more diverse set %of of  taxa but individual taxa would not be as numerically dominant as in the more %uniform uniform  environment inhabited by the plankton. At the Order level, enriched %bacterial bacterial  OTUs tended to have members in that were enriched in both the %plankton plankton  and the biofilm suggesting the phylogenetic coherence of lifestyle is %not not  captured at the level of Order. It should be noted however that taxonomic %annotations annotations  in reference databases and therefore environmental sequence %collections collections  show little equivalency in phylogenetic breadth between groups at %the the  same taxonomic rank \citep{Schloss_2011}. Unfortunately, at higher %taxonomic taxonomic  resolution (e.g. Genus-level), groups did not possess a sufficient %number number  of OTUs to evaluate coherence between taxonomic annotation and %lifestyle. lifestyle.  Similar to the richness results, we found the %shape shape  of rank abundance distributions between biofilm and planktonic libraries %in in  our data (Figure~\ref{fig:rank_abund_shape} to be in contrast to that %reported reported  by \citet{22237539} although this may be an artifact of each study's %different different  source communities (as %discussed discussed  above). Carbon amendments did not affect algal library membership %and and  structure to the same degree as it affected bacterial library composition. %As As  expected, bacterial OTUs enriched in the high C amended mesocosm (C:P = %500) 500)  include OTUs in classic copiotroph families such as \textit{Vibrionacaea} %and and  \textit{Pseudomonadaceae}. Interestingly, the one OTU depleted in the high %C C  treatments is annotated as being in the HTCC2188 order of the %\textit{Gammaproteobacteria}. \textit{Gammaproteobacteria}.  HTCC stands for 'high throughput culture %collection' collection'  and is a prefix for strains cultured under low nutrient conditions %\citep{Cho_2004, \citep{Cho_2004,  Connon_2002}. %\subsection{Conclusion}  %In summary this study shows mechanistic links between large scale community