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\section*{Acknowledgments}  COPO is funded by a UK Biotechnology and Bioscience Research Council (BBSRC) Bioinformatics and Biological Resources Fund (BBR) grant: BB/L021390/1, BB/L024055/1, BB/L021390/1 [BB/L024055/1,  BB/L024071/1, BB/L024101/1. BB/L024101/1].