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The constant interaction of processes, such as weathering, atmospheric deposition, nutrient leaching, and biological cycling, results in the formation of vertical and horizontal nutrient gradients within the soil (Fig. 1)\cite{Giehl2014}.   Which root traits for efficient nutrient uptake?  \cite{York2013a} theoretical framework of classification of root phenes in base of their function, distinguing between exploration and exploitation, and cost. Complement with different possibilities for limitations to root growth in \cite{Postma2014a}.  \section{Root development}  The formation of LRs is a complex and highly coordinated process that depends on the interplay of hormones, where auxin acts as a central component. Indeed, the deficiency of many nutrients has been reported to affect the levels of one or more hormones in plants (for review, see Rubio et al., 2009)\cite{Giehl2014}.