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\section{Introduction}
Ecosystem subsidies, the transfer of resources across ecosystem boundries (cite) are an important part of organic matter cycling in aquatic systems. Although the reciprical transfer of resources between aquatic and terrestrial systems is common (cite), the majority of lentic and lotic ecosystems rely heavily on terrestrial organic matter inputs to support their metabolism and secondary productionMarcarelli \cite{Marcarelli_2011}. Organic matter subsidies from terrestrial to aquatic ecosystems are dominated by detrital plant material either as dissolved (DOC) or particulate (POC) organic matter carbon. Although DOC and POC inputs are variable across systems, they can represent a substantial augmentation of autochthonous organic matter production \cite{Hodkinson_1975} \cite{GASITH_1976} \cite{wetzel_1984} \cite{WETZEL_1995} \cite{Webster_1997} \cite{kobyashietal2011}
\cite{Mehring_2014}meheringetal2014}. \cite{Mehring_2014}.
%Allochthonous organic matter inputs are an important basal resource in many aquatic systems. Terrestrial leaf litter inputs determine the biogeochemistry and trophic dynamics of small streams \cite{websterandbenfield1986,wallaceetal1997,wallaceetal1999} and recently it has been shown that temperate lake food webs depend on dissolved allochthonous organic matter \ref{paceetal#####}. However, lakes also receive substantial inputs of terrestrial leaf litter \ref{hodkinson1975, reed1979, wallaceandbenfield1986, franceandpeters1995, kobayashietal2011}, and the presence of litter can alter lentic nutrient cycling \ref{wetzel} and community structure \ref{}.