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\section{Introduction}  Ecosystem subsidies, the movement of resources across ecosystem boundries \cite{Polis_1997}, are an important part of organic matter cycling in aquatic systems. Although the reciprical transfer of resources between aquatic and terrestrial systems is common \cite{Nakano_2001}, the majority of lentic and lotic ecosystems rely heavily on terrestrial organic matter inputs to support their metabolism and secondary production \cite{Marcarelli_2011}. Organic matter subsidies from terrestrial to aquatic ecosystems are dominated by detrital plant material either as dissolved (DOC) or particulate (POC) organic carbon, which can represent a substantial augmentation of autochthonous organic matter production \cite{Hodkinson_1975,GASITH_1976,wetzel_1984,WETZEL_1995,Webster_1997,Kobayashi_2011,Mehring_2014}.   During the process of leaf litter decomposition in aquatic systems, the mass of the leaf can supply distinct supplements to the aquatic system \cite{Gessner_1999}. Up to 30\% of the initial mass of leaves can be leached as DOC {CITE}, (CITE),  although large initial DOC fluxes from dried leaves may be an artifact of the drying process {CITE}. (CITE).  This supply of DOC is an important component of aquatic organic matter budgets \cite{McDowell_1976,Karlsson_2007}} \cite{McDowell_1976,Karlsson_2007}  and has been shown to alter the abundance \cite{Bott_1984,Fey_2015} and function \cite{MCCONNELL_1968,Bernhardt_2002} of aquatic microbial communities. Furthermore, DOC subsidies processed through the microbial loop can support metazoan production \cite{Hall_1998,Wilcox_2005,Fey_2015b}.