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At the Arctic, self-organization for all groups of phyto and zooplankton species in all zones, were high or very high. Only the low emergences of diatoms, cyanobacteria and green algae permits that these photosynthetic organisms reach very high levels of complexity and autopoiesis. This situation continuing in NH in spite of planktonic diatoms and cyanobacteria reached the fair category of emergence. It means, the feature of this two types of lakes is their regularity.  In a similar way that with limiting nutrient subsystem, whenthe  gradient Ar-T reachesat  the NL point point,  the dynamics of emergence and self-organization varies in considerable level. Here, the complexity of almost all variables were maximum due the balance in self-organization and emergence. Only, chlorophiceas, benthic detritus and fishes in planktonic and benthic zones have regularity for all annual cycle. In contrast to the NL, the biomass subsystem in the tropic reflects very low complexity due the very high self-organization of the living taxa. Only planktonic and piscivorous fishes have very high and high complexity, respectively.