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\subsubsection{Complexity in Biomass Subsystem}  At the Arctic Arctic,  self-organization for all groups of phyto and zooplankton species in all zones, are were  high or very high. Only the low emergences of Diatoms, Cyanobacteria diatoms, cyanobacteria  and Green Algae green algae  permits that these photosynthetic organisms reach very high levels of complexity and autopoiesis. This situation continuing in NH in spite ofthe  planktonic diatom diatoms  and cyanobacteria reach reached the  fair category of emergence. It means, the feature of this two types of lakes is your their  regularity. In a similar way that with limiting nutrient subsystem subsystem,  when the gradient reach Ar-T reaches at the  NL point the  dynamics of emergence and self-organization varies in considerable level. Here, the complexity of almost all variables were maximum due the balance in self-organization and emergence reached. emergence.  Only, Chlorophiceas, Benthic chlorophiceas, benthic  detritus and Fishes fishes  in Planktonic planktonic  and Benthic Zone benthic zones  have regularity for  all annual cycle cycle.  In contrast to the NL, the biomass subsystem in the tropic reflects very low complexity due the very high self-organization of the living taxa. Only planktonic and piscivorous fishes reached have  very high and high complexity, respectively. In terms of autopoiesis, the autonomy of the biomass at the tropic demonstrates that this subsystem could be affected in major proportions in case of climatic change events. From PCA ordination, it can be seen that photosynthetic taxa of planktonic and benthonic zone are more emergent and macrophytes. emergent.  Consumers primaries and secondaries of the plankton in addition with clorophiceas and BDt are more self-organized and self-organized.  Piscivorous and Planktivorous fishes are more complex.The last passively due to…  \subsubsection{Complexity in Latitudinal Gradient}  \subsection{Autopoiesis}  In terms of autopoiesis, the autonomy of the biomass at the tropic demonstrates that this subsystem could be affected in major proportions in case of climatic change events.  \subsection{Homeostasis}