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\subsection{Complexity in ecology and measures of complexity in ecology}  McArthur (1955) were the first researcher to recognize the important role of Shannon’s entropy in ecological studies.  Ricotta:..."there have been few attempts to quantify the statistical complexity of a given species assemblage  in ecology; Demetrius et al. (2004) examine the question from an evolutionary biology perspective; Demetrius and Manke (2005) present methods which can be applied to ecological networks (e.g., food webs); Juh´asz-Nagy (1976, 1984, 1993); Juh´ asz-Nagy and Podani (1983) was the first to develop a coherent information-theoretical framework for summarizing various aspects of spatial complexity in plant communities as a function of scale. (Ricotta and Anand, 2004): A measure of statistical complexity for plant communities based on Juh´ asz- Nagy’s model.