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other measures... transfer entropy, Tsallis entropy...Rao`s quadratic entropy index (function dbFD from package FD in R; also check ade4) with presence-absence data, that it is simply the mean distance between the occurring species. (Inpect the scaling by the possible maximum, because it coudlbe meaningless for abundance data).  http://www.uoguelph.ca/~manand/Papers/RicottaEt06.pdf: "Shannon’s entropy, fail to adequately capture the structural complexity of   an ecological community. The contributions of spatial complexity to community structure   are however quantifiable if we consider, for example, the degree of spatial co-occurrence   between species. The larger and more intricate these correlations, the more structurally   complex the community. We suggest that Juhasz-Nagy information-theoretical functions ´   offer an adequate basis for the measurement of structural complexity""  http://chaos.aip.org/resource/1/chaoeh/v18/i4/p043106_s1  plasticity, memory, and cognition as special cases of emergence, self-organization, and complexity.