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\subsection*{Statistical analysis}  Statistical analysis was performed using R (version 3.2.0)\cite{R} except data analysis of electrophysiological recordings (GraphPad, San Diego, CA, USA).  For comparison between two genotypes, we used Welch's t-test.  Mann-Whitney U test was performed for data with non-normal distribution.  For comparison among three and more genotypes, we used one-way analysis of variance (ANOVA) followed by Tukey HSD post hoc test.   Welch's one-way analysis of variance (ANOVA) followed by Games-Howell post hoc test was performed for data with unequal variance.  We used "car"\cite{car} and "userfriendlyscience"\cite{user} R packages for Levene's test for homogeneity of variances and Welch's one-way ANOVA test, respectively.      
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