ACKNOWLEDGEMENTS
We acknowledge support for this research from NSF Dimension of Biodiversity grant 1738041. This research was supported financially by the Perennial Agriculture Project, a joint project between The Land Institute and The Malone Family Land Preservation Foundation. We gratefully acknowledge financial support from the Foundation for Food and Agriculture Research (FFAR) Fellows program. We thank Eric Cassetta for assisting with all of the field work. We are thankful to Andrew Read and Peter Innes for assistance with code to generate the phylogeny and map figures, respectively.