Acknowledgements
The authors thank the staff of the School of Biological Sciences for their technical support and advice as well as the late Ms Tsz Chun So for sample collection in the field. The computations were performed using research computing facilities offered by Information Technology Services, at the University of Hong Kong. This study was partially supported by the Research Grants Council of Hong Kong (GRF 17113221) and by the NSFC/RGC Joint Research Scheme (Grant No. N_HKU791/22)