Author contributions
Q.K. conceived and designed the study and revised the manuscript. Y.Z. and X.Z. provided the plant materials. H.W. participated in data analysis, writing and revising the manuscript. X.W. participated in sampling and sequencing. C.M. and M.W. participated in data analysis. Y.L., Y.F., and X.G. contributed to the preparation of plant samples. All authors read and approved the final manuscript.