Acknowledgements
We sincerely appreciate the assistance in turion collection from Dr. Yang Li, Dr. Lei Yang and Dr. Tian Lv from Wuhan University. The present study was financially supported by the National Natural Science Foundation of China (31800299), the general financial grant from the China Postdoctoral Science Foundation (2017M622184) and the Startup Foundation for Introduced Talents of Qingdao Agricultural University (663/1116023).