Acknowledgments
This study was funded by National Natural Science Funds(32070178),Shandong Provincial Modern Agricultural Industry and Technology System (SDAIT-08-06), Shandong Province Agriculture Major Application Technology Innovation Project(SD2019XM007),Talent Engineering Projects (ts201511069; W03020496), Agricultural Science and Technology Innovation Project of Shandong Academy of Agricultural Sciences(CXGC2021A12 and CXGC2021A15) ,Independent training innovation team of Jinan (2019GXRC025) and National Key Research and Development Program of China (2017YFC1702700).