ACKNOWLEDGEMENTS
The research was supported by the National Key R&D Program, Ministry of Science and Technology, China [2017YFD0300103], the National Natural Science Foundation of China (31771719), and National High Technology Research and Development Program of China (2014AA10A605). Rothamsted Research receives strategic funding from the Biotechnological and Biological Sciences Research Council of the UK. MP acknowledges funding from the Designing Future Wheat Strategic Programme (BB/P016855/1).