Running title: Hawaiian arthropods and habitat fragmentation

Emma Steigerwald, Museum of Vertebrate Zoology, University of California, Berkeley; Department of Environmental Science, Policy, and Management, University of California, Berkeley

Judith Paetsch, Department of Biogeography, University of Trier, Trier, Germany

Dana Drück, Department of Biogeography, University of Trier, Trier, Germany

Jana Fritsch, Department of Biogeography, University of Trier, Trier, Germany

Marie Klaka, Department of Biogeography, University of Trier, Trier, Germany

Susan R. Kennedy, Department of Biogeography, University of Trier, Trier, Germany

Rosemary G. Gillespie, Department of Environmental Science, Policy, and Management, University of California, Berkeley

Henrik Krehenwinkel, Department of Biogeography, University of Trier, Trier, Germany*

* corresponding author: krehenwinkel@uni-trier.de