AUTHOR CONTRIBUTIONS
The fieldwork was carried out by HJ. DS, SLL and AH. AKN and HJ developed the pedigree. DS carried out genetic assignment. DS and SLL performed the statistical analyses and interpreted the results with input from AKN, DG, SM and HJ. DS drafted the initial version of the manuscript with input from SLL, DG, AKN, SM and HJ. All authors contributed to later versions of the manuscript, and all authors gave final approval for publication.