Funding
This work was supported by National Natural Science Foundation of China (No.22078128, 21938004), the 111 Project (No. 111-2-06), the Fundamental Research Funds for the Central Universities (No. JUSRP221032), Postgraduate Research & Practice Innovation Program of Jiangsu Province KYCX21-2028, National First-Class Discipline Program of Light Industry Technology and Engineering (LITE2018-24), the Opening Project of the Key Laboratory of Industrial Biotechnology, Ministry of Education (KLIBKF201802), British Council 527429894 and BBSRC BB/T013176/1.