Physiological analysis
After measuring the dry weight of the upper part and roots of plants that were treated for 15 days (Figure 1A ), a decrease in the upper part dry weight (DW) was observed in salinity plus B-deprivation (Comb (-)) treatment compared to control plants. The salinity (NaCl) and Comb (-) treatments were lower and statistically different upper part values from plants treated with B excess (B (+)), which had the highest upper part total dry mass. Additionally, NaCl and Comb (-) treated plants presented lower root DW than the control plants. The analysis of relative water content showed no differences of any of the treatments with the control plants. However, while B (-) and B (+) showed no significant differences, when they were combined with salinity, Comb (-) were significantly lower than Comb (+). The Leaf per Mass Area (LMA) data (Figure 1B ) showed that only the salinity leaves were significantly different from the control plants, providing higher values.
We analyzed stomatal conductance (Figure 2 ) every 3-4 days from the beginning to the end of the experiment. The control and B (-) plants showed the highest stomatal conductance that were maintained during all the time of the experiment. Plants treated with B (+) maintained similar values than control until the 3rd day of treatment showing from that day a significant decrease that remain stable from 10th to 13th day of treatment. Salinity (NaCl), Comb (-), and Comb (+) treatments had the lowest stomatal conductance that decreased at first day of treatment, about half that of the control plants, being the lowest those plants treated with NaCl. During the experiment, the stomatal conductance of B (+) treated plants reached similar stomatal conductance to the salinity-related treatments at 10th day of treatment.
The mineral analysis of leaves (Figure 3A ) showed a clear division between salinity-related treatments and non-salinity treatments along the horizontal axis and between B (-) and B (+) treatments (including Comb (-) and Comb (+) respectively) along the vertical axis, as summarized by a Principal Component Analysis (PCA). The interactions between Na and K, as well as between B and Mn, were negative. However, Cu and Zn demonstrated a positive interaction with NaCl and Comb (-) treatments, but a negative interaction with treatments related to excess B (B (+) and Comb (+)). Figure 3B-D illustrates individual plots of B, Na, and K concentrations as mass per DW represented as box plots. The control plants had around 50 mg mg kg-1 DW of B, while NaCl, B (-), and Comb (-) treatments had lower concentrations, being Comb (-) leaves the one that presented the lowest concentration of B among all treated plants with around 10 mg kg-1 DW. Alternatively, B (+) treated plants had the highest amount of B at 109 mg kg-1 DW, doubling the control B concentration, followed by Comb (+) at 87 mg kg-1 DW. The three salinity-related treatments (NaCl and the two combinations) had 3.0-3.5 g of Na per 100 g DW, while the rest of the plants had marginal concentrations. In contrast, K levels were lower these salinity treatments, along with B (-), which also had a reduction in K presence compared to control plant leaves, but it was still higher than the salinity treatments.