Variation in frond area was correlated with natural levels of light availability (F1,32 = 16.99, p=0.0002, β= -9.663, R2=0.35), with plants growing in more heavily shaded sites with thicker canopy cover expressing larger fronds (Fig. 2A). Variation in root length was correlated with water nutrient levels (both dissolved and total Nitrogen and Phosphorus), with plants growing longer roots in lower nutrient conditions (Fig. 2B). The variable with the most explanatory power was total Phosphorus content (F1,32 = 27.59, p<0.0001, β= -0.10, R2=0.46). PH failed to explain additional variation in these traits.