Author Contributions
H.J., W.S., X.W.,and C.S. conceptualized the initial study; H.J.,M.Z., and F. Y. were involved in the experimental layout and carried out experiment and analyzed experiment results; H.J., X.L., J.J., Y.Z., Y.W., and T.J. were analyzed experiment results; Q.W., and M.Z. performed the subcellular localization experiments; H.J. drafted original manuscript and provided funding; C.S. provided funding and edited the manuscript.