ACKNOWLEDGEMENTS
We thank the staff of the Guilin Botanical Garden for their contribution in the field. This study was supported by the National Natural Science Foundation of China (No.31870514; 32170492); Guangxi Natural Science Foundation (No.2019GXNSFD- A245021); Doctoral startup fund of Guangxi Normal University for Nationalities (No. 2021BS002); The fourth batch of characteristic discipline construction projects in Ethnic Colleges and universities approved by the Department of education of Guangxi Zhuang Autonomous Region (Ethnic Ecology). The basic ability enhancement program for young and middle-aged teachers of Guangxi university (2020KY20017). The funders had no role in study design, data collection and analysis, decision to publish, or preparation of the manuscript.