Acknowledgements
This study was funded by the National Natural Science Foundation of China (41701081, 42061013), Guizhou Basic Research Project (QKHJC-ZK[2022]146, QKHJC[2020]1Y118), Young Talents Project of Guizhou Education Department (KY[2017]119) and Guizhou Science and Technology Project ([2018 ]4007(006)). We would like to thank Ling Luo and Qiumei Zhao for their help with field soil sampling and measurement.