Author contributions

L. wrote the manuscript. W.L., M.C., H.T., and G.W. performed data analysis. C.Z., M.C., H.T., and G.W. performed experiments. Y.M., Y.H., M.C., amd Y.M. were responsible for collecting the samples. X.Z., B.Y., and M.P. revised the manuscript. X.Z. and B.Y. designed and supervised the study.