ACKNOWLEDGEMENTS
We thank Prof. Xiang-Zong Meng (Shanghai Normal University) and Prof. Yi-Ming Wang (Nanjing Agricultural University) for providingArabidopsis mutants. We also thank Dr. Da-Cheng Wang, Yun Guan and Fan-Qi Kong (Nanjing Agricultural University) for their technical support. This work was supported by the National Natural Science Foundation of China (32272617, 31972322, 31701829), the Science and Technology Project of Jiangsu Province (BE2021364, BE2020408), the Fundamental Research Funds for the Central Universities (KYZZ2022001), the Opening Project of Key Construction Laboratory of Probiotics in Jiangsu Province (JSYSZJ2019003).