Data analysis
For assessing plant growth parameters, enzyme activity, and gene expression analysis, statistical significance was used for analysis of variance (One-way ANOVA) in SPSS 22.0 (SPSS Inc., Chicago, IL, USA). Duncan’s post-test (P <0.05) was used for multiple comparisons. The results were expressed as the mean ± SE . In all figures, different capital letters indicate significant differences between same treatments, under the different water condition, whereas the lowercase letters indicate significant differences between different treatments seedlings under the same water condition. The experiment was carried out with three biological replicates, and each experimental index was measured by at least four replicates.