Where Bi,c is the niche breadth of speciesi with respect to climate variable c , and p is the proportion of that species’ SDM found in state k of the climatic variable (e.g., within each 1°C increment of temperature or 250 mm increment of precipitation). To facilitate comparison across species and different niche variables, we calculated a standardized niche breadth,\(\hat{B}\)i,c , bound between zero and one, by dividing niche breadth by the number of possible states (n ), corrected for the finite number of states: