ACKNOWLEDGMENTS
We thank Natural England for permits and landowners for access to populations; Ferhat Celep and Judita Kochjarová for assistance during fieldwork collection. This research was supported by the Graduate Campus office at the University of Zurich through a GRC-Travel Grant and the Forschungskredit Candoc (grant no. K-74202-05-01) to EM-C, by the Swiss Government Excellence Scholarship (grant no. 2018.0475) to EM-C, by the Swiss National Science Foundation (grant no. 310030_185251) to JdV, and (grant nos. 3100-061674.00/1 and 31003A_175556) to EC.