Acknowledgement
This work was supported by the National Key Research and Development Program of China (2018YFD0501506);
The Natural Science Foundation of Shandong Province (ZR2019YQ17);
The Higher Education Support Program of Youth Innovation and Technology of Shandong Province, China(2019KJF022);
The China Agriculture Research System (CARS-42-19);
Funds of Shandong “Double Top” Program (SYL2017YSTD11).