Author contribution
F.Q. participated in designing experiments, carried out the experiments in this study, prepared the first draft of this manuscript. Z.L. and Q.W. conceived of the study, participated in its design. A.L., Y.J., Z.L., H.Z., D.N., Q.L., H.W., S.L. performed a part of experiments. Q.L., J.L., S.L. participated in experimental design and consulted on the study. All authors approved the final manuscript.