Acknowledgements:
We are grateful to William E. (Ned) Friedman for kindly sharing plant material and lab facilities and to the staff of the Arnold Arboretum of Harvard University for continuous support. This work was funded by the National Science Foundation IOS 1456845 research grant to N.M.H. Z.H. was awarded with a research internship program from the DaRin Butz Foundation at the Arnold Arboretum of Harvard University. JML is a ComFuturo researcher at the IHSM La Mayora, funded by FGCSIC, a RTI2018-102222-A-I00 grant from the Spanish Ministry of Science and Universities, and a LINKB20067 from CSIC.