Acknowledgments
We thank three anonymous reviewers and the editor for constructive comments and suggestions to make this paper greatly improved. This study is supported by the Second Tibetan Plateau Scientific Expedition and Research (STEP) program (2019QZKK0503), the National Science Foundation of China (41877048 and 41825016), US Department of Energy, Office of Science, Genomic Science Program under Award Numbers DE-SC0004601 and DE-SC0010715, the NSF LTER program, and the Office of the Vice President for Research at the University of Oklahoma. The authors declare no conflict of interest.